Wood Formation In Trees

Trees are a major component of the biosphere and have played an important part in the world's history and culture. With the modern challenges of global warming and dwindling fossil fuel reserves, trees, and in particular their wood, can provide solutions. Unfortunately, too little is known about the biology of these plants, due largely to a lack of appropriate techniques. In recognition of this, Wood Formation in Trees presents a variety of detailed techniques and protocols for the study of the cell and molecular biology of wood formation in trees. Internationally recognized experts, most of whom are the researchers who developed the techniques, speak with authority in this volume, and also provide first-hand tips and trade secrets to help the uninitiated master the techniques. The techniques reflect a hierarchical approach to the study of the developmental biology of wood formation: anatomical, biochemical and molecular-genetic. Trees are a tremendous but vastly under-appreciated natural resource. In an age where the natural product is so often modified to suit modern tastes and industrial processes, it is essential to understand how the natural product is made. The techniques in this book provide that essential information about the process of wood formation in trees.

Trees are a major component of the biosphere and have played an important part in the world's history and culture. With the modern challenges of global warming and dwindling fossil fuel reserves, trees, and in particular their wood, can provide solutions. Unfortunately, too little is known about the biology of these plants, due largely to a lack of

The book is a fundamental reference source on reaction wood for wood scientists and technologists, plant biologists, silviculturists, forest ecologists, and anyone involved in the growing of trees and the processing of wood. It brings together our current understanding of all aspects of reaction wood, and is the first book to discuss both compression wood and tension wood. Trees produce reaction wood to maintain the vertical orientation of their stems and the optimum angle of each branch. They achieve this by laying down fibre cell walls in which differences in physical and chemical structure from those of normal fibres are expressed as differential stresses across the stem or branch. This process, while of obvious value for the survival of the tree, causes serious problems for the utilisation of timber. Timber derived from trees containing significant amounts of reaction wood is subject to dimensional instability on drying, causing twisting, bending and splitting. It is also difficult to work as timber, and for the pulp and paper industry the cost of removing the increased amount of lignin in compression wood is substantial. This has both practical and economic consequences for industry. Understanding the factors controlling reaction wood formation and its effect on wood structure is therefore fundamental to our understanding of the adaptation of trees to their environment and to the sustainable use of wood. The topics covered include: -Morphology, anatomy and ultrastructure of reaction wood -Cell-wall polymers in reaction wood and their biosynthesis -Changes in tree proteomes during reaction wood formation -The biomechanical action and biological functions of reaction wood - Physical and mechanical properties of reaction wood from the scale of cell walls to planks -The detection and characterisation of compression wood -Effects of reaction wood on the performance of wood and wood-based products - Commercial implications of reaction wood and the influence of forest management on its formation

The Formation of Wood in Forest Trees covers the proceedings of the second symposium held under the auspices of the Maria Moors Cabot Foundation for Botanical Research, conducted in Harvard Forest in Petersham, Massachusetts on April 15-19, 1963. The book focuses on the aspects of tree growth, such as the chemistry and submicroscopic morphology of wood and the effects of the environment on growth. The selection first offers information on the evolution of cambium in geologic time; a model for cell production by the cambium of conifers; and structure and development of the bark in dicotyledons. The text then ponders on the aspects of ultrastructure of phloem, stem structure in arborescent monocotyledons, and structure and formation of the cell wall in xylem. The publication takes a look at the general chemistry of cell walls and distribution of the chemical constituents across the walls and ultraviolet and fluorescence optics of lignified cell walls. The text also examines the role of endogenous hormones in cambial activity and xylem differentiation; indirect effects of environment on wood formation; and influence of external pressure on the differentiation of cells and tissues cultured in vitro. The selection is a dependable reference for readers interested in the formation of wood in forest trees.

With today’s ever growing economic and ecological problems, wood as a raw material takes on increasing significance as the most important renewable source of energy and as industrial feedstock for numerous products. Its chemical and anatomical structure and the excellent properties that result allow wood to be processed into the most diverse products; from logs to furniture and veneers, and from wood chippings to wooden composites and paper. The aim of this book is to review advances in research on the cellular aspects of cambial growth and wood formation in trees over recent decades. The book is divided into two major parts. The first part covers the basic process of wood biosynthesis, focusing on five major steps that are involved in this process: cell division, cell expansion, secondary cell wall formation, programmed cell death and heartwood formation. The second part of the book deals with the regulation of wood formation by endogenous and exogenous factors. On the endogenous level the emphasis is placed on two aspects: control of wood formation by phytohormones and by molecular mechanisms. Apart from endogenous factors, various exogenous effects (such as climate factors) are involved in wood formation. Due to modern microscopic as well as molecular techniques, the understanding of wood formation has progressed significantly over the last decade. Emphasizing the cellular aspects, this book first gives an overview of the basic process of wood formation, before it focuses on factors involved in the regulation of this process.

"First published in 1945 by Collins"--Copyright page.