Nearshore Marine Paleoclimatic Regions Increasing Zoogeographic Provinciality Molluscan Extinctions And Paleoshorelines California

Approximately 3000 middle and late Cenozoic nearshore marine molluscan taxa from western California are assigned to six time periods, spanning ~25 m.y. In this interdisciplinary study, western California is palinspastically restored for each of the time periods by backsliding and back-rotating large fault blocks or crustal units. Marine fossil assemblages are assigned to nearshore paleoclimatic regions or water masses within palinspastically restored California. In addition, this volume reveals positive feedback mechanisms between paleolatitudinal changes in sea-surface paleotemperature gradients and changes in the diversity of marine mollusks along the California coast through time; defines "equable" based effective temperatures; and analyzes extinction rates among macroinvertebrate marine taxa from coastal California and the possible causes of these extinctions. The late Paleogene to Neogene faunas reflect an increase in faunal diversity related to strengthened temperature gradients, greater extremes in sea-surface temperatures, reduction in temperateness, and the development of an embayed California coastline.

“A masterful accomplishment—Allen, Pondella and Horn have assembled a talented team of experts who produce authoritative, up-to-date accounts. This book will be used as the primary text in many fish biology courses and as a valuable reference elsewhere. Here is a wealth of data waiting to be mined by legions of graduate students as they generate the new ideas that will motivate marine ecology for years.”—Peter Sale, Editor of Coral Reef Fishes: Dynamics and Diversity in a Complex Ecosystem "A copiously illustrated and comprehensive interpretation of the past, present, and future state of over 500 species of fishes in Californian waters. A compilation of virtually all the many important studies on the ecology of California marine fishes."—Bruce B. Collette, National Marine Fisheries Service and co-author of The Diversity of Fishes

Phylogeography of California examines the evolution of a variety of taxaÑancient and recent, native and migratoryÑto elucidate evolutionary events both major and minor that shaped the distribution, radiation, and speciation of the biota of California. The book also interprets evolutionary history in a geological context and reviews new and emerging phylogeographic patterns. Focusing on a region that is defined by physical and political boundaries, Kristina A. Schierenbeck provides a phylogeographic survey of CaliforniaÕs diverse flora and fauna according to their major organismal groups. Life history and ecological characteristics, which play prominent roles in the various outcomes for respective clades, are also considered throughout the work. Supporting scholars and researchers who study evolutionary diversification, the book analyzes research that helps assess one of the major challenges in phylogeographic studies: understanding changes in population structures shaped by geological and geographical processes. California is one of only twenty-five acknowledged biological hotspots worldwide, and the phylogeographic history of the state can be extrapolated to study other regions in western North America. Further consideration is given to implications for conservation, recommendations concerning the biogeographic provinces that roughly define the state of California, and predictions related to climate change.

In a three part study, we analyzed: (1) population-level response of the coachwhip snake, (Masticophis flagellum) and striped racer (M. lateralis) to fragmentation of scrub habitats in coastal southern California using site occupancy models and presence/apparent absence data; (2) space-use behavior of coachwhip snakes and racers in fragmented and continuous habitats of coastal southern California using data from radio-telemetry; and (3) phylogeography and population genetic variation throughout the range of both species with emphasis on coachwhip populations in southern California using mitochondrial DNA sequence and RFLP data. Part 1 revealed disproportionate loss of coachwhip populations from fragmented areas. Unlike the racer, the coachwhip tended to be absent from sites embedded in isolated landscapes, even at large spatial scales. The two species showed large differences in rates of local extinction and probability of detection, with coachwhip populations exhibiting variable rates of detection and greater likelihood of local extinction. Habitat composition influenced site occupancy. Coachwhip snakes preferred sites with open sage scrub and grassland. Racers preferred scrub dominated sites. In Part 2 coachwhip snakes exhibited great plasticity in spacing and movement patterns between sites, showing large differences in movement response to changes in local conditions and amount of available space. In sympatry, coachwhip snakes and racers showed large differences in home range size, rate of travel, and movement distance, with coachwhip snakes ranging more widely than racers. Differences in habitat use fit well with occupancy patterns. Coachwhip snakes utilized both scrub and grassland, and racers utilized largely only scrub. In Part 3 phylogenetic analyses recovered significant structure in both species, with seven well-supported and geographically coherent clades recovered from coachwhip populations sampled throughout the range of the species. High density sampling within southern California recovered four cryptic coachwhip lineages in contact and organized geographically by bioregion. Collectively, results suggest the greater vulnerability of the coachwhip to land-use change is related to differences in habitat use, movement, and rates of population turnover. Unless the few remaining large, contiguous landscapes of grassland and open scrubland are incorporated into regional conservation planning, long-term persistence of coachwhip lineages is unlikely in the future landscape of California.

With all the recent advances in molecular and evolutionary biology, one could almost wonder why we need the fossil record. Molecular sequence data can resolve taxonomic relationships, experiments with fruit flies demonstrate evolution and development in real time, and field studies of Galapagos finches have provided the strongest evidence for natural selection ever measured in the wild. What, then, can fossils teach us that living organisms cannot? Evolutionary Patterns demonstrates the rich variety of clues to evolution that can be gleaned from the fossil record. Chief among these are the major trends and anomalies in species development revealed only by "deep time," such as periodic mass extinctions and species that remain unchanged in form for millions of years. Contributors explore modes of development, the tempo of speciation and extinction, and macroevolutionary patterns and trends. The result is an important contribution to paleobiology and evolutionary biology, and a spirited defense of the fossil record as a crucial tool for understanding evolution and development. The contributors are Ann F. Budd, Efstathia Bura, Leo W. Buss, Mike Foote, Jörn Geister, Stephen Jay Gould, Eckart Hâkansson, Jean-Georges Harmelin, Lee-Ann C. Hayek, Jeremy B. C. Jackson, Kenneth G. Johnson, Nancy Knowlton, Scott Lidgard, Frank K. McKinney, Daniel W. McShea, Ross H. Nehm, Beth Okamura, John M. Pandolfi, Paul D. Taylor, and Erik Thomsen.