The Great Ordovician Biodiversification Event

Two of the greatest evolutionary events in the history of life on Earth occurred during Early Paleozoic time. The first was the Cambrian explosion of skeletonized marine animals about 540 million years ago. The second was the "Great Ordovician Biodiversification Event," which is the focus of this book. During the 46-million-year Ordovician Period (489–443 m.y.), a bewildering array of adaptive radiations of "Paleozoic- and Modern-type" biotas appeared in marine habitats, the first animals (arthropods) walked on land, and the first non-vascular bryophyte-like plants (based on their cryptospore record) colonized terrestrial areas with damp environments. This book represents a compilation by a large team of Ordovician specialists from around the world, who have enthusiastically cooperated to produce this first globally orientated, internationally sponsored IGCP (International Geological Correlation Program) project on Ordovician biotas. The major part is an assembly of genus- and species-level diversity data for the many Ordovician fossil groups. The book also presents an evaluation of how each group diversified through Ordovician time, with assessments of patterns of change and rates of origination and extinction. As such, it will become the standard work and data source for biotic studies on the Ordovician Period.

The graptolites constitute one of the geologically most useful taxonomic groups of fossils for dating rock successions, understanding paleobiogeography and reconstructing plate tectonic configurations in the Lower Palaeozoic. Graptolites were largely planktic, marine organisms, and as one of the first groups that explored the expanses of the world’s oceans are vital for understanding Palaeozoic ecology. They are the best and often the only fossil group for dating Lower Palaeozoic rock successions precisely. Thousands of taxa have been described from all over the planet and are used for a wide variety of geological and palaeontological (biological) research topics. The recent recognition of the modern pterobranch Rhabdopleura as a living benthic graptolite enables a much better understanding and interpretation of the fossil Graptolithina. In the decades since the latest edition of the Graptolite Treatise, the enormous increase of knowledge on this group of organisms has never been synthesised in a compelling and coherent way, and information is scattered in scientific publications and difficult to sort through. This volume provides an up-to-date insight into research on graptolites. Such research has advanced considerably with the use of new methods of investigation and documentation. SEM investigation and research on ultrastructure of the tubaria has made it possible to compare extant and extinct taxa in much more detail. Cladistic interpretation of graptolite taxonomy and evolution has advanced the understanding of this group of organisms considerably in the last two decades, and has highlighted their importance in our understanding of evolutionary processes. This book will show graptolites, including their modern, living relatives, in a quite new and fascinating light, and will demonstrate the impact that the group has had on the evolution of the modern marine ecosystem. This book is aimed not only at earth scientists but also at biologists, ecologists and oceanographers. It is a readable and comprehensible volume for students at the MSc level, while remaining accessible to undergraduates and non-specialists seeking up-to-date information about this fascinating topic in palaeobiology.

This book provides a complete Phanerozoic story of palaeogeography, using new and detailed full-colour maps, to link surface and deep-Earth processes.

The Early Palaeozoic was a critical interval in the evolution of marine life on our planet. Through a window of some 120 million years, the Cambrian Explosion, Great Ordovician Biodiversification Event, End Ordovician Extinction and the subsequent Silurian Recovery established a steep trajectory of increasing marine biodiversity that started in the Late Proterozoic and continued into the Devonian. Biogeography is a key property of virtually all organisms; their distributional ranges, mapped out on a mosaic of changing palaeogeography, have played important roles in modulating the diversity and evolution of marine life. This Memoir first introduces the content, some of the concepts involved in describing and interpreting palaeobiogeography, and the changing Early Palaeozoic geography is illustrated through a series of time slices. The subsequent 26 chapters, compiled by some 130 authors from over 20 countries, describe and analyse distributional and in many cases diversity data for all the major biotic groups plotted on current palaeogeographic maps. Nearly a quarter of a century after the publication of the ‘Green Book’ (Geological Society, London, Memoir12, edited by McKerrow and Scotese), improved stratigraphic and taxonomic data together with more accurate, digitized palaeogeographic maps, have confirmed the central role of palaeobiogeography in understanding the evolution of Early Palaeozoic ecosystems and their biotas.

Special Publication 485 About 40 million years after the Cambrian Explosion, the Great Ordovician Biodiversification Event (GOBE) represents a second and dramatic burst in marine biodiversity, with major changes in the structure of ecosystems and the progressive replacement of the distinctive Cambrian Evolutionary Fauna by the Paleozoic Evolutionary Fauna. However, the GOBE is not a single, worldwide, short-term event, but rather the complex sum of successive diversifications occurring in distinct taxonomic groups, trophic guilds and regions. This book focuses on the Late Ordovician Tafilalt Biota, Anti-Atlas Morocco, which provides a snapshot of the GOBE in high-latitude regions of the Southern Hemisphere. A series of contributions explore different aspects of the Tafilalt Biota, including its geological setting, the international fossil trade in this area and a series of detailed systematic contributions describing many new taxa of marine invertebrates. This volume represents a significant contribution to the understanding of the Tafilalt Biota and its significance to the GOBE.

World-class palaeontologists and biologists summarise the state-of-the-art on fish evolution and development.

This volume addresses major evolutionary changes that took place during the Ediacaran and the Paleozoic. These include discussions on the nature of Ediacaran ecosystems, as well as the ichnologic signature of evolutionary radiations, such as the Cambrian explosion and the Great Ordovician biodiversification event, the invasion of the land, and the end-Permian mass extinction. This volume set provides innovative reviews of the major evolutionary events in the history of life from an ichnologic perspective. Because the long temporal range of trace fossils has been commonly emphasized, biogenic structures have been traditionally overlooked in macroevolution. However, comparisons of ichnofaunas through geologic time do reveal the changing ecology of organism-substrate interactions. The use of trace fossils in evolutionary paleoecology represents a new trend that is opening a window for our understanding of major evolutionary radiations and mass extinctions. Trace fossils provide crucial evidence for the recognition of spatial and temporal patterns and processes associated with paleoecologic breakthroughs.

This 3-volume handbook brings together contributions by the world ́s leading specialists that reflect the broad spectrum of modern palaeoanthropology, thus presenting an indispensable resource for professionals and students alike. Vol. 1 reviews principles, methods, and approaches, recounting recent advances and state-of-the-art knowledge in phylogenetic analysis, palaeoecology and evolutionary theory and philosophy. Vol. 2 examines primate origins, evolution, behaviour, and adaptive variety, emphasizing integration of fossil data with contemporary knowledge of the behaviour and ecology of living primates in natural environments. Vol. 3 deals with fossil and molecular evidence for the evolution of Homo sapiens and its fossil relatives.